where we came from. The science of paleoanthropology, which also tries to create a
narrative about how humans came to be, is deeply technical. Paleoantropology is the
and it is the paleoanthropologist?s job to identify and explain these changes. In this
Homo sapiens are the only living representative of the family Hominidae. The
brains. So all humans are hominids, but not all hominids could be called human. Next
binocular vision, mobile fingers and toes with flat nails instead of claws, a shortened
are unusual for mammals, humans are even more unusual for primates. We are
essentially elaborated African apes. We share almost 99 percent of our genetic material
with chimpanzees. Yet we have several traits that are very different.
Two legged walking, or bipedalism seems to be one of the earliest of the major
hominine characteristics to have evolved. To accommodate this strange position, we
have developed a specialized pelvis, hip and leg muscles, and an S-shaped vertebral
column. Because these changes can be documented in fossil bone, bipedalism is seen
as the defining trait of the sub family Homininae.
Much of the human ability to make and use tools and other objects stem from
braincase volume of between 79.3 and 91.5 cubic inches. In the course of human
evolution the size of the brain has more than tripled. The increase in brain size may be
became increasingly numerous and sophisticated. It is likely that the increase in
human brain size took place as part of a complex interrelationship that included the
elaboration of tool use and tool making, as well as other learned skills which permitted
which may reflect a pattern of the different sexes in our early ancestors. The bones
suggest that females may have been 3 to 4 ft in height and about 60 to 70 lb. in weight,
while males may have been somewhat more than about 5 ft tall, weighing about 150 lb.
patterns of behavior in early hominine social groups. This extreme difference between
sexes appears to disappear gradually sometime after a million years ago.
The third major trend in hominine development is the gradual decrease in the
size of the face and teeth. All the great apes are equipped with large, tusklike canine
teeth that project well beyond the level of the other teeth. The earliest hominine
remains possess canines that project slightly, but those of all later hominines show a
marked reduction in size. Also, the chewing teeth, the premolars and molars, have
decreased in size over time. Associated with these changes is a gradual reduction in the
size of the face and jaws. In early hominines, the face was large and positioned in front
of the braincase. As the teeth became smaller and the brain expanded, the face became
smaller and its position changed. Thus, the relatively small face of modern humans is
located below, rather than in front of, the large, expanded braincase.
Evidence of immediate relatives of the human species begins about five million
years ago with the Australopithecus genus and leads in to the primitive Homo genus to
modern humans. The nature of the human?s evolution before that is uncertain, but
million years ago, primitive apelike animals were widely distributed on the African and
later on the Eurasian continents. Although many fossil bones and teeth have been
of these fossil apes, known as Sivapithecus, appears to share many features with the
living Asian great ape and the orangutan, whose direct ancestor it may well be. None
leading to the hominid family generally. But they do help paint a picture of what early
human relatives could have been like.
The convincing fossil evidence for human evolution begins with
Australopithecus. Fossils of this genus have been discovered in a number of sites in
fossils show them existing about 3.9 million years ago, and the genus flourished until it
seemed to have become extinct about 1.5 million years ago. All the australopithecines
were efficiently bipedal and thus indisputable hominines. In details of their teeth, jaws,
and brain size, however, they differ enough among themselves to warrant division into
four species: A. afarensis, A. africanus, A. robustus, and A. boisei.
between 3 and 3.9 million years ago. Found in the Afar region of what is now Ethiopia
and in Tanzania, A. afarensis had a brain size a little larger than those of chimpanzees.
Some of the species possessed canine teeth somewhat more projecting than those of
later hominines. No tools of any kind have been found with A. afarensis fossils.
Between about 2.5 and 3 million years ago, A. afarensis apparently evolved into
a later australopithecine, A. africanus. Known primarily from sites in southern Africa,
A. africanus possessed a brain similar to that of its predecessor. However, although
the size of the chewing teeth remained large, the canines, instead of projecting, grew
only to the level of the other teeth. As with A. afarensis, no stone tools have been
found in association with A. africanus fossils.
By about 2.6 million years ago, the fossil evidence reveals the presence of at
least two, and perhaps as many as four, separate species of hominines. An evolutionary
split seems to have occurred in the hominine line, with one group evolving toward the
genus Homo, and finally to modern humans, and the others developing into
australopithecine species that eventually became extinct. The australopithecine species
that eventually became extinct includes the robust australopithecines, A. robustus, that
lived in southern Africa, and A. boisei, found only in eastern Africa. The robust
australopithecines represent an unusual adaptation because their principal difference
from other australopithecines lies in the large size of their chewing teeth, jaws, and jaw
muscles. The robust australopithecines became extinct about 1.5 million years ago.
Although scientists do not agree, many believe that after the evolutionary split
that led to robust australopithecines, A. africanus evolved into the genus Homo. This
was a species called Homo habilis, or “handy man.” Appearing about 2.5 million years
ago, the new hominid probably didn’t look terribly different from its predecessors, but it
had a somewhat larger brain. And, perhaps as a result of some mental connection other
termites out of their mounds (something modern chimps still do). But Homo habilis
deliberately hammered on rocks to crack and flake them into useful shapes. The tools
average, was less than 5 ft. tall and weighed under 100 lbs., and it could hardly have
left over from predators’ kills. Even other scavengers like hyenas, jackals and the such
were stronger and tougher than early humans. But Homo habilis presumably had the
tools to butcher leftovers quickly and get back to safety.
Their adaptations to the rigors of prehistoric African life enabled members of
the Homo habilis species to survive for 500,000 years or more, and at least one group
of them apparently evolved, around 2 million years ago.
Around this time, East African mammals adapted to drier more open grassland
conditions. It was about this time that the new form of human emerged in Africa, a
hominid with a much larger brain, excellent vision, and limbs and hips fully adapted to
an upright posture. Paleoanthropologists call this hominid Homo Erectus, a human
much taller than its diminutive predecessors, standing on average five feet six inches
tall, with hands capable of precision gripping and many kinds of tool-making. The
skull is more rounded than those of earlier hominids, but still had a sloping forehead
and retreating brow ridges. Homo Erectus was more numerous and more adaptable
than Homo habilis, and, on present evidence, was a much longer lived species.
Archaeological sites for this species appear at higher, cooler elevations in southern,
eastern, and northern Africa. Homo erectus may have been a skilled big game hunter,
capable of organizing quite elaborate hunting and foraging expeditions, and using
multipurpose axes and cleaving tools.
Like all hunters and foragers, Homo Erectus had probably learned to live with
natural fires and was not afraid of them. In time, the new hominid may have made a
habit of conserving fire, taking advantage of smoldering tree stumps ignited by
of all, the making of fire. Perhaps as early as 1.5 million years ago, Homo erectus may
offers not only warmth, but protection against predators and an easy way of hunting
game, even insects and rodents. The toxins from many common vegetable foods can
Homo erectus was a much larger species than its predecessors meaning that the
newcomers needed larger quantities of food to satisfy higher metabolic rates. This
meant they had to range over much larger hunting territories perhaps moving into more
them as a weapon that would enable them to operate safely away from trees, and to
and live in far cooler environments. It may be no coincidence that the earliest human
Somewhere between 200,000 and 300,000 years ago, Homo erectus evolved
during this period and exactly when it happened. In fact, certain fossils from this period
are classified as late Homo erectus by some scientists and as early Homo sapiens by
others, depending on the scientists belief in what happened.
Even though they are in the same genus and species as modern humans, these
early Homo sapiens do not have identical physical traits to modern humans. New fossil
evidence suggests that modern man, sometimes called Homo sapiens sapiens (a
sub-species of Homo sapiens), first appeared more than 90,000 years ago. There is
some disagreement among scientists on whether the hominine fossil record shows a
continuous evolutionary development from the first appearance of Homo sapiens to
modern humans. This disagreement has especially focused on the place of Neandertals
evolution. The Neanderthals (named for the Neander Valley in Germany, where one of
100,000 years ago until about 35,000 to 40,000 years ago, when they disappeared from
the fossil record. Fossils of additional varieties of early Homo sapiens have been
discovered in other parts of the Eurasia.
The dispute over the Neanderthals also involves the question of the evolutionary
term race is not possible (because modern humans show continuous variation from one
geographic area to another), widely separate human populations are marked by a
number of physical differences. Most of these differences represent adaptations to
spread of Homo erectus sometime after a million years ago. In their view, human
development since Homo erectus has been one continuous, in-position evolution,
meaning, local populations have remained, changing in appearance over time. What
they are trying to say is that the peopling of the world, the spreading of humans, has not
changed since Homo Erectus. The Neanderthals and other early Homo sapiens are
seen as descending from Homo erectus and are ancestral to modern humans.
Other scientists view racial differentiation as a relatively recent phenomenon. In
their opinion, the features of the Neanderthals which are a low, sloping forehead, large
brow ridge, and a large face without a chin are too primitive for them to be considered
the ancestors of modern humans. They place the Neanderthals on a side branch of the
modern humans can be found in southern Africa or the Middle East. Evolving perhaps
90,000 to 200,000 years ago, these humans then spread to all parts of the world,
supplanting the local, earlier Homo sapiens populations. In addition to some
fragmentary fossil finds from southern Africa, support for this theory comes from
comparisons of mitochondrial DNA, a DNA form inherited only from the mother,
suggest that humans derived from a single generation in southern Africa or
southeastern Asia. Because of the tracing through the material line, this work has come
to be called the ?Eve? hypothesis. Its results are not accepted by most scientists, who
consider the human race to be much older.
Whatever the outcome of this scientific disagreement, the evidence shows that
early Homo sapiens groups were highly efficient at exploiting the sometimes harsh
climates of Ice Age Europe. Further, for the first time in human evolution, hominines
began to bury their dead deliberately, the bodies sometimes being accompanied by
stone tools, by animal bones, and even by flowers.
Although the evolutionary appearance of modern peoples did not dramatically
change the basic pattern of adaptation that had characterized the earlier stages of
human history, some innovations did take place. In addition to the first appearance of
during this time that human language originated, a development that would have had
profound implications for all aspects of human activity. About 10,000 years ago, one of
human history that eventually led to civilization.
The next evolutionary stage, is modern humans as we know them. Early Homo
sapiens developed the traits that were discussed earlier, and man as we know it, came
to exist. But how do we know all these things occurred, and why? It all comes from
several very important steps in paleoanthropology. The earliest human skeleton ever
found was discovered in 1974 in a remote region of Ethiopia, a very well preserved A.
afarensis fossil. Nicknamed Lucy, paleoanthropologists have found out periods of
millions of years through her, and have used that knowledge to speculate on other
discoveries. Footprints, bones, teeth, etc. are all used to help piece together the puzzle.
Human evolution may have reached a dead end, foreseeable for a while at least.
Despite the enormous changes that we have wreaked on our environment, major
evolutionary changes in humans will not occur in the distant future. Scientists dismiss
the idea that the species is ?going somewhere? under natural selection and then
describe how most successful species are stable through their geological lifetimes.
populations, there is little chance for a new different human species.
Modern understanding of human evolution rests on known fossils, but the
picture is far from complete. Only future fossil discoveries will enable scientists to fill
many of the blanks in the present picture of human evolution. Employing sophisticated
technological devices as well as the accumulated knowledge of the patterns of
geological deposition, anthropologists are now able to pinpoint the most promising
locations for fossil hunting more accurately. In the years ahead this will result in an
enormous increase in the understanding of human biological history.
and Hudson, 1990.
We Come From? Where Are We Going?? Science 14 Jan. 1994: 181-183
Gibbons, Ann, ?When It Comes to Evolution, Humans Are in the Slow Class.? Science
31 March. 1995: 1907-1908
?Human Evolution.? Microsoft Encarta. 1996 ed. [CD-ROM]
Lemonick, Michael ?New Thinking on Human Evolution? Time 14 March 1994: 81-87